Neuroscience PhD Program (NDP)

NDP PhD Program banner


Cheryl Stucky, PhD


Cheryl Stucky, PhD

Professor of Cell Biology, Neurobiology, & Anatomy
Director, Neuroscience Doctoral Program


The Neuroscience Doctoral Program (NDP) at MCW is devoted to training the next generation of neuroscience researchers. A diverse group of neuroscientists with an impressive record of pre-doctoral training, research and outside funding, form the core faculty for this doctoral training program. Throughout the course of study, students will interact with a large number of faculty, students and postdoctoral trainees engaged in cutting-edge neuroscience research. The goal is to provide each student with the basics of modern neuroscience and then allow customization of a program of course work that best meets his/her needs.
(414) 955-8373


The following departments support the Neuroscience Doctoral Program in Biomedical Sciences through faculty mentorship and funding. Newly-enrolled students will have the ability to choose faculty from one of the departments below to rotate with throughout their first year in the program. Students are then matched into one of the labs that they rotated in at the conclusion of their first year, and transition from being a NDP student to a student of their faculty mentor’s department.

Biophysics (PhD) 

MCW’s Biophysics PhD program is home to the National Biomedical Electron Paramagnetic Resonance (EPR) Center and features two areas of primary research: Molecular Biophysics and Magnetic Resonance Imaging (MRI). As a Biophysics student, you will be able to choose from a variety of research topics, ranging from the application of new technologies in solving biological problems to cognitive neuroscience to structural and functional dynamics of proteins. If you are interested in conducting biophysics research, we encourage you to apply directly to the program, or by applying to either our Interdisciplinary Program (IDP) in Biomedical Research or the Neuroscience Doctoral Program (NDP).

Cell & Developmental Biology (PhD) 

The Cell & Developmental Biology PhD program at MCW is defined by its research strengths in cellular and molecular mechanisms of organ development, stem cell biology and its impact on regenerative medicine and neuroscience. This program is home to MCW's Regenerative Medicine and Stem Cell Biology program, which focuses on the study of normal development and disease pathways of the liver, heart and nervous system. Some of the noteworthy research breakthroughs within Cell & Developmental Biology include the characterization of novel mutations underlying Axenfeld-Reiger Syndrome as well as the role of TRPA1 channels in nociception. If you are interested in participating in cell and developmental biology research, you can seek admission into MCW’s Graduate School by applying directly to the program through either the Interdisciplinary Program (IDP) or Neuroscience Doctoral Program (NDP).

Biochemistry (PhD)

The Biochemistry PhD program at MCW will expose you to state-of-the-art facilities and instruments for 3D structure determination of proteins and protein-drug complexes by X-ray crystallography, fluorescence microscopy and nuclear magnetic resonance (NMR) spectroscopy. The research areas of our faculty are broad and include, but are not limited to, cell & development biology, structural biology and drug discovery. If you are interested in conducting biochemistry research at MCW, we invite you to apply through either our Interdisciplinary Program (IDP) or Neuroscience Doctoral Program (NDP).

Pharmacology & Toxicology (PhD) 

The Pharmacology & Toxicology PhD program at MCW emphasizes research in mechanistic studies of drug actions/signaling molecules and is home to the Drug Discovery Center, which focuses on the translation of new discoveries into therapies that can be used to improve human health. If you are interested in pharmacology and toxicology research, you can seek admission into MCW’s Graduate School by applying through either the Interdisciplinary Program (IDP) or the Neuroscience Doctoral Program (NDP).

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IDP-Biophysics Graduate ProgramAbout the Neuroscience Doctoral Program (NDP)


The Neuroscience Doctoral Program (NDP) of the Medical College of Wisconsin is an exciting, rigorous and innovative PhD research program that provides highly motivated students from around the world with first-class training in Neuroscience Research.

Our 20 year history offers state-of-the-art research training in Neuroscience domains that range from molecular and cellular signaling to human brain imaging, in over 30 labs within 18 departments and centers, all with strong national funding. Close connections between clinical and basic faculty within our academic medical institution presents many opportunities for students to conduct clinically-relevant research on diseases of the nervous system.

Our highly selective program admits on average 5 students per year, allowing the students close interactions and mentoring by our excellent faculty mentors and teachers. The program is scholarly and challenging but student-centered and flexible, and our goal is to help our students develop the knowledge, laboratory skills, critical thinking and communications skills that are essential for a successful career in Neuroscience.

Current Students


Neuroscience Doctoral Program Students

Current  Neuroscience Doctoral Program Students

NDP students and Cheryl Stucky, PhD, spend a day volunteering at the Hunger Task Force in Franklin, WI. Harvesting corn for 5 hours was hard work but lots of fun and very rewarding!

"I like MCW because of the collaborative platform for research, the atmosphere and the interdisciplinary research environment where the knowledge can be transferred to a clinical application."

Seung-Yi Lee
2nd year NDP student

"Seeking to further understand the underlying mechanisms of the visual system and its interaction with the brain truly excites me about neuroscience."

Jonathon Young
3rd year NDP student

"Courses in computer science and neuroscience led me to join a lab that marries neurobiology and computer science. This made the NDP and Joe Carroll’s lab at MCW an easy choice."

Alexander Salmon
4th year NDP student


Thiago ArzuaThiago N. Arzua

Thesis Lab: Xiaowen Bai, MD, PhD, Department of Anesthesiology
Research Interests: Neuroscience involves the possibility to further develop almost every field of research. When there is a need to push the boundaries of neuroscience, technology, methodologies, society and arguably even philosophy are all pushed with it. It is incredibly exciting to be able to tackle questions that can create such extensive themes.
Year Entered MCW: 2016
Undergraduate Education: University of South Florida, Tampa, FL, Chemistry with a minor in French


Eric ClarkEric Clark

Thesis Lab: Brian Link, PhD, Department of Cell Biology, Neurobiology & Anatomy
Research Interests: Current research projects include studying mutations in genes that encode proteins involved in mitochondria trafficking and dynamics, as well as extracellular matrix proteins. The overall goal is to understand how these factors contribute to Neurodegenerative Disease.
Year Entered MCW: 2011
Undergraduate Education: Albion College, Albion, Michigan, Major: biology/psychology, neuroscience concentration

Matt DillardMatthew Dillard

Thesis Lab: Matt Hodges, PhD, Department of Physiology
Research Interests: I believe that the most exciting part of Neuroscience is the breadth of the field. There are a wide variety of problems to be addressed in the field which provides ample opportunity for the advancement of knowledge in the field and the application of new discoveries for the betterment of humanity. I also find the interdisciplinary nature of Neuroscience to be exciting. One problem in the field can be addressed in numerous different ways by individuals with widely differing backgrounds. The synthesis of knowledge and methodologies from various disciplines makes Neuroscience research exciting to me.
Year Entered MCW: 2016
Undergraduate Education: Eureka College, Eureka, Ill, Neuroscience

Ethan DuwellEthan Duwell

Thesis Lab: Edgar A. DeYoe, PhD, Department of Biophysics
Research Interests: My research interests include visual attention, conscious awareness, and conscious experience. I am currently pursuing these interests in the DeYoe lab, where we study visual attention both behaviorally and neurophysiologically via functional magnetic resonance imaging (fMRI).
Year Entered MCW: 2015
Undergraduate Education: St. Olaf College (Northfield, MN), BA, Chemistry and Philosophy

Heather HeitkotterHeather Heitkotter

Thesis Lab: Joseph Carroll, PhD, Cell Biology, Neurobiology & Anatomy
1st year student
Research Interests: My general focus is neuroscience; I enjoy investigating the underlying mechanisms of behavior and neurological function. I have a particular interest in neurological development and related disorders, which I hope to explore via cellular activities and phenotypic expression.
Year Entered MCW: 2017
Undergraduate Education: Lake Forest College; Neuroscience and Biology

Ryan HillmerRyan Hillmer

Thesis Lab: Brian Link, PhD, Department of Cell Biology, Neurobiology & Anatomy
Research Interests: An interest in neurodevelopment and neurodegenerative disease led me to join a lab investigating the molecular mechanisms governing embryonic development. I am currently investigating the interactions between Notch and Hippo signaling early in development using the zebrafish model system. These interactions are probed in the zebrafish eye by measuring signaling activity of the Notch and Hippo pathways in fish with altered levels of Hippo and Notch signaling, respectively.
Year Entered MCW: 2012
Undergraduate Education: Carroll University, BS

Ben OBrien HokensonBen O'Brien Hokanson

1st year student
Research Interests: My research interests focus around neurodegenerative disease because of the complexity found in these conditions and the lack of suitable treatments currently. I am interested in pursuing pharmacological treatments for these diseases as I progress with my studies. 
Year Entered MCW: 2017
Undergraduate Education: University of Wisconsin Madison, Madison, WI, Neurobiology

Seung-Yi LeeSeung-Yi Lee

Thesis Lab: Matt Budde, PhD, Department of Neurosurgery
Research Interests: I studied specific ion effects on hydrophobic hydration of model polymers that are reminiscent to proteins at Penn State. One of the applications of my studies was for understanding neurodegenerative diseases which led me to neuroscience.
Undergraduate Education:  Catholic University of Korea, Seoul, South Korea; Chemistry, English Literature/ Education
Year Entered MCW: 2016
Graduate Education: Pennsylvania State University, State College, PA; Bioanalytical Chemistry

Xiaojie LiuXiaojie Liu

Thesis Lab: Qing-Song Liu, PhD, Department of Pharmacology & Toxicology
Research Interests: My research focuses on understanding the mechanism of cocaine addiction. Addiction involves the changes of the dopamine reward system, which consists of dopamine projections from the ventral tegmental area (VTA) to different brain regions. My interest is in identifying the drug-induced changes in the reward circuit, and then investigating whether enhancing and blocking the identified change affects drug-induced addictive behaviors.
Year Entered MCW: 2015
Undergraduate Education: Shanxi Medical University, Taiyuan, China; Neurophysiology

Sean McGarrySean McGarry

Thesis Lab: Peter LaViolette, PhD, Department of Radiology
Research Interests: My research interests involve structural imaging in brain cancer.
Year Entered MCW: 2015
Undergraduate Education: Milwaukee School of Engineering (Milwaukee, WI), BS, Biomedical Engineering

Francie MoehringFrancie Moehring

Thesis Lab: Cheryl Stucky, PhD, Department of Cell Biology, Neurobiology & Anatomy
Research Interests: My research focuses on the idea that while sensory neurons have long been known to mediate touch and pain transduction, they are highly likely not alone and epidermal keratinocytes are likely the initial first responders to tactile stimuli. Therefore I am trying to dissect the cellular mechanism by which keratinocytes communicate with sensory nerve terminals.
Year Entered MCW: 2013
Undergraduate Education: University of Wisconsin – Milwaukee, BS, Biology, Chemistry

Natalie NawarawongNatalie N. Nawarawong

Thesis Lab: Christopher Olsen, PhD, Department of Pharmacology & Toxicology
Research Interests: My research will focus on expanding our understanding of the neural populations and pathways that underlie both drug and non-drug rewards. By using transgenic mouse models, I will be able to investigate the cellular and molecular mechanisms of addiction.
Year Entered MCW: 2012
Undergraduate Education: Vassar College

Savannah NeuSavannah Neu

1st year student
Research Interests: Great potential has been shown in recent years for a connection between inflammation in the Central Nervous System (CNS) and the microbiome of the human gut.  My current research interest involves studying how a person's environment (ie. diet, infection) can modulate the inflammatory response of autoimmune neurodegenerative diseases like Multiple Sclerosis (MS).
Year Entered MCW: 2017
Undergraduate Education: Milwaukee School of Engineering, Major: BioMolecular Engineering; Minors: Mathematics, Physics, Chemistry

Ashley ReynoldsAshley Reynolds

Thesis Lab: Cheryl Stucky, PhD, Department of Cell Biology, Neurobiology & Anatomy
Research Interests: The inflammasome is responsible for production of the pain-inducing cytokine IL-1β. Sensory neurons release IL-1β and directly respond to IL-1β because sensory neurons express its receptor IL-1R1. Therefore, it is likely that the inflammasome is responsible for the release of IL-1β from sensory neurons and involved in pain. Thus, I am studying the role of the sensory neuron inflammasome in postoperative pain.
Year Entered MCW: 2014
Undergraduate Education: Marquette University, BS, Biochemistry and Molecular Biology

Jose RosasJose Rosas

1st year student
Research Interests:
Year Entered MCW: 2017
Undergraduate Education:

Ben SajdakBenjamin Sajdak

Thesis Lab: Joseph Carroll, PhD, Cell Biology, Neurobiology & Anatomy
Research Interests: I am using non-invasive retinal imaging techniques like optical coherence tomography and adaptive optics scanning light ophthalmoscopy to study retinal disease. I work with several species, but most often the ground squirrel. I love studying this model because it is a cone dominant rodent, and undergoes incredible metabolic changes with annual hibernation.
Year Entered MCW: 2014
Undergraduate Education: University of Wisconsin Oshkosh, Oshkosh, WI; Major: Biology, Minor: Neuroscience.

Alexander SalmonAlex Salmon

Thesis Lab: Joseph Carroll, PhD, Cell Biology, Neurobiology & Anatomy
Research Interests: Software, hardware, and biological solutions to improve adaptive optics scanning light ophthalmoscopy (AOSLO) and optical coherence tomography (OCT) imaging in animals. Developing animal models of retinal disease. Developing contrast agents for OCT.
Year Entered MCW: 2014
Undergraduate Education: University of Wisconsin-Madison, Madison, WI, Neurobiology

Stephanie SantarriagaStephanie Santarriaga

Thesis Lab: Matt Scaglione, PhD, Department of Biochemistry
Research Interests: My research project is to understand mechanisms by which Dictyostelium resists protein aggregation. A hallmark of aging and age-related diseases is the loss of proteostasis, leading to an increase in protein aggregation. The nine polyglutamine diseases are caused by the expansion of CAG trinucleotide repeat within the coding region of specific genes. CAG codes for the amino acid glutamine, resulting in the translation of a polyglutamine-expanded, aggregation-prone protein. We identified Dictyostelium discoideum as a model organism that normally encodes for long polyglutamine tracts and is highly resistant to polyglutamine aggregation.
Year Entered MCW: 2013
Undergraduate Education: University of Minnesota, Duluth, MN, Biochemistry, Cell Biology, and Molecular Biology

Rebecca SlickRebecca Slick

1st year student
Research Interests: I’ve always been excited about neuroscience because it is utilized in a wide variety of fields. I’m interested in doing research that investigates behavior as well as neurological function in a translational setting.
Year Entered MCW: 2017
Undergraduate Education: University of Wisconsin - Oshkosh

Jiaqing TongJiaqing Tong

Thesis Lab: Jeffery R. Binder, MD, Department of Neurology
Research Interests: Exploring how the neurons code the information when people are involved in certain activities interests me.
Undergraduate Education: Shanxi Medical University, Taiyuan, Shanxi, Clinical Lab Technician
Year Entered MCW: 2016
Graduate Education: Shanxi Medical University, Taiyuan, Shanxi, Physiology

Jonathon YoungJonathon B. Young

Thesis Lab: Iris Kassem, MD, PhD, Cell Biology, Neurobiology & Anatomy
Research Interests: Inflammation and fibrosis following lensectomy in a juvenile rabbit model – The goal of this project is to ultimately improve the outcomes of cataract surgery performed in infants.
Year Entered MCW: 2015
Undergraduate Education: Marquette University, Milwaukee, WI; Biomedical Science

Hanmeng ZhagHanmeng Zhang

Thesis Lab: Daniel Lipinski, MSc, DPhil, Ocular Gene Therapy Lab
Research Interests:  The nervous system plays a leading role in the regulation of body functions. It is related to multiple disciplines and they are inseparable. I hope that after the collection of broad neuroscience knowledge I will connect them with sports. I believe that “exercise is medicine” and hope that we will find more possibilities which exercises will bring to us.
Year Entered MCW: 2016
Undergraduate Education: Beijing Sport University, Beijing, China; Human Sport Science

Program Overview and Structure



Program Highlights

  • No teaching requirement, may have teaching options in advanced years with PI approval
  • Students are encouraged to attend and present at Society for Neuroscience, Keystone and Gordon conferences. Travel funds available.
  • Students can take courses at other institutes (UWM, MU).
  • Rotations in 1st year allow exploration of research interests, mentoring styles and lab dynamics experiencing neuroscience from genes to behavior
  • Coursework beyond 1st semester is flexible & tailored to each student’s interest
  • Small graduate class size allow ample faculty-student interaction

Program Summary

During the first year, students spend their mornings taking fundamental courses in molecular, cellular and systems neuroscience. In the afternoons, students begin doing hands-on research through 9 week rotations in 3-4 laboratories of their choosing. By the end of the first year, they select a lab and mentor for their thesis work. In years 2-3, students take advanced courses that are tailored to their research interests and thesis goals. Students also take a comprehensive qualifier exam that includes writing an NIH-style innovative research proposal.

Throughout all years, students have the opportunity to present at regional, national and even international neuroscience meetings, and often receive competitive internal and external travel awards. Students have the opportunity to gain teaching experience by assisting in graduate and medical neuroscience courses at MCW. On average, students defend their dissertation work in 51/2 years from beginning the program.

Our NDP alumni advance to exciting careers that include postdoctoral fellowships at premier research institutes, pharmaceutical companies and faculty positions at research and teaching colleges.

First Year

  • Attend weekly Journal Club to discuss primary literature with faculty member.
  • Rotate through labs of Neuroscience faculty to gain hands-on lab experience and explore the variety of research opportunities.
  • Select a research mentor and join the graduate program of the mentor's department after completion of the first year.
  • Course work includes Biochemistry of the Cell, Molecular and Cellular Biology, Fundamentals of Neuroscience and Mechanisms of Cellular Signaling or Molecular Genetics.

Second Year

  • Complete a mock grant proposal that serves as the qualifying exam.
  • Strong emphasis is placed on completion of dissertation research on a state-of-the-art problem of current importance in neuroscience.
  • Choose from elective courses such as Biology of Vision, Physiological Genomics, Cellular Signal Transduction, Advanced Systems Neuroscience, Advanced Cell Biology, Cellular Molecular Neurobiology and Classical and Molecular Genetics 

Third Year and beyond

  • During the subsequent years of study, emphasis will be placed on advanced training in selected areas of neuroscience in conjunction with dissertation research in a problem of current importance in the neurosciences.
  • Choose from elective courses such as Biology of Vision, Physiological Genomics, Cellular Signal Transduction, Advanced Systems Neuroscience, Advanced Cell Biology, Cellular Molecular Neurobiology and Classical and Molecular Genetic


Listed below are the required and elective courses most often taken by our NDP students. There are many additional courses offered by other departments at the Medical College of Wisconsin that can be taken by students in the Interdisciplinary Neuroscience Program depending upon their research interests. In addition, graduate courses offered at Marquette University or University of Wisconsin Milwaukee can be taken for credit to satisfy some elective requirements; more information is available from the MCW Graduate School of Biomedical Sciences.

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  Biochemistry of the Cell

Course Number: 16202

Number of Credits: 4


This interdisciplinary course provides students with a solid foundation in the understanding of the structure and function of proteins. This knowledge is then applied to proteins involved in various metabolic pathways to understand the manner in which these pathways are organized and controlled. The material is presented primarily in lecture format, interspersed with occasional discussion sessions.

  Techniques in Molecular and Cellular Biology

Course Number: 16242
Number of Credits: 2


The objective for the Techniques course is to provide a theoretical and practical foundation underlying a number of the most common experimental techniques required for biomedical research. The information presented in this course will introduce procedures and experimental strategies that are commonly used in biomedical research projects, and will facilitate students’ comprehension of the scientific literature even if they don’t use the techniques in their own research. The lecture materials present the theory behind each technique, the practical limitations of each techniques, and the types of questions that each technique addresses, with emphasis on how each can be applied to generate new insight into biomedical research questions.

  Molecular and Cellular Biology

Course Number:16244

Number of Credits: 4


This interdisciplinary course provides students with a solid foundation in the areas of gene expression and cell biology. The material is presented primarily in lecture format, but a significant number of discussion sections are also included.

  Fundamentals of Neuroscience

Course Number:12211
Number of Credits: 4

This course follows a multidisciplinary approach to current knowledge about the structural and functional properties of the nervous system. The mechanisms of the nervous system are described at the molecular, cellular, and multi-cellular levels. The course includes both lectures and laboratory sessions. The purpose of this course is to introduce PhD students to the anatomy and function of the human nervous system.

  Mechanisms of Cellular Signaling

Course Number: 16250

Number of Credits: 4


This interdisciplinary course provides first-year graduate students with a foundation in cellular signal transduction. The course has three sections; in the first, students learn the basic building blocks of signaling, including ligands, receptors and adaptor proteins; in the second section, students learn about representative signaling cascades; and in the third section, students consider signaling in the context of cellular or tissue biology. The material is presented in lectures, primary paper discussions and in open-ended discussion sessions.

  Classical and Molecular Genetics

Course Number: 16252

Number of Credits: 4


This interdisciplinary course provides students with a foundation in classical and molecular genetics, model systems genetics, the replication, repair and recombination of the genetic material, developmental biology, cancer, and genomics. The material is presented primarily in lecture format, but a significant number of discussion sections are also included.

  Biology of Vision

Course Number: 31257

Number of Credits: 3


This lecture/discussion course explores the functional organization and development of the visual system as revealed by the use of a variety of anatomical, cell biological, genetic, physiological and behavioral methods. It is designed for students who wish to gain a basic understanding of the biological basis for vision and to share in the excitement of the latest developments in this field. Topics include: Development of the eye and visual system, fundamental principles of regulated gene expression, the cell biology of the photoreceptors and retina, phototransduction and neural processing in the retina, functional architecture of retina and visual system, the anatomy, physiology and perceptual significance of parallel pathways.

  Physiological Genomes

Course Number: 08230

Number of Credits: 5


This course will cover topics in Physiological Genomics at an advanced level emphasizing the tools and techniques that are available to investigators exploring the relationship between genotype and phenotype. Material will be selected to emphasize high throughput screening and Bioinformatics techniques. Specific examples of applications of physiological genomics to important research problems will be discussed. Students will acquire the expertise required to develop a research proposal and will participate in a mock study section to witness the process by which grants are reviewed.

  Essential Physiological Genomics

Course Number: 08229

Number of Credits: 2


This course covers genome sequence, functional genomic analysis, genome and gene manipulation, and grant writing. The students will learn about the latest advances in the field of physiological genomics, how to apply genomic approaches to study complex physiological problems and how to develop a grant proposal.

  Cellular Signal Transduction

Course Number: 07224

Number of Credits: 3


This course provides an in-depth presentation of mechanisms of cellular signaling at a level designed for doctoral students in the biomedical sciences. The emphasis is on receptors, second messenger systems, G proteins and signal transduction.

  Advanced Systems Neuroscience

Course Number: 12221

Number of Credits: 3


This course covers seven selected areas in systems neuroscience, including: neuronal information processing and control systems, cerebral hemodynamics, metabolism and neuronal activity, sensory systems, motor systems, attentional systems, learning and memory and motivational systems. Some lectures introducing fundamental concepts and current research topics are presented but learning occurs primarily through readings and discussions.

  Advanced Cell Biology

Course Number: 31250

Number of Credits: 3


Lectures and readings in the renewal, differentiation, communication, adhesion, secretion, motility, gene activity, and mitochondrial dynamics of eukaryotic cells.

  Cellular and Molecular Neurobiology

Course Number: 12237

Number of Credits: 3


Readings and discussion in cellular, molecular, and developmental neurobiology. Among the topics covered in this course are ion channels and the ionic basis of potentials; mechanisms of synaptic transmission; neurotransmitter receptors and their receptors; sensory signal transduction and neural development.

  Journal Club

Course Number: 12298

Number of Credits: 1


Weekly readings will be selected from both contemporary and historical literature in neuroscience. Informal discussions will include participation from both neuroscience faculty and students

Applying to the Program


Dr. Cheryl Stucky with NDP students

Start Your Application Now

First year students meet with Cheryl Stucky, PhD, to discuss a recent article at the weekly Neuroscience Journal Club meeting.

We are delighted that you are considering MCW for your graduate education. In an effort to help make your application process as efficient as possible, this webpage contains all the information necessary to complete your application. Below you will find descriptions of the academic and technical admission requirements and a list of the items that make up a completed application.


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  Online Application

Due by January 15 (for Fall term enrollment)


Application Date


December 15, 2017


December 16, 2017 - January 15, 2018



Applications received on or by December 15, 2017 will be waived of the admission fee. It is in the candidate’s best interest to have all materials in by this date. Applications received December 16, 2017 or later will be charged the normal admission fee of $50. In special circumstances of financial need, a waiver for reduced fee may be requested. Applications received by December 15, 2017 will be given top priority in review. However applications received after that date will continue to be considered until all slots for the NDP program are filled.


Complete a degree seeking online application, with no fee by December 15, 2017. Standard application fee waivers will still be offered after the priority application date. Visit the MCW Graduate School admissions webpage for more information.



Send official transcripts from all current or previously attended undergraduate, graduate and medical institutions directly to the MCW Graduate School. Transcripts in the hands of prospective students are not accepted as "official" unless they are submitted in their original envelope with signed seals intact. A grade point average of 3.0 or higher is ideal. Electronic transcripts may be accepted from institutions that use either eSCRIP-SAFE, National Clearinghouse and/or Parchment transcript delivery companies.


GRE Results

Send an official Graduate Record Exam (GRE) test score report directly from ETS to the MCW Graduate School (Institution Code 1519). Applicants who have earned a PhD, MD, JD or other terminal degree may request an exemption to the GRE test score submission by submitting a written request to Applicants are encourage to explore whether they may be eligible for the GRE Fee Reduction Program.


Letters of Recommendation

Send three letters of recommendation. Recommenders should submit their letter of recommendation along with an accompanying recommendation form (PDF) via scan-and-email to or by mail to: 


MCW Graduate School
8701 Watertown Plank Rd.
Milwaukee, WI 53226  



The Test of English as a Foreign Language (TOEFL) is required of all applicants who did not receive a bachelor’s or master’s degree from a U.S.-based, non-online degree granting program. Please arrange for your TOEFL scores to be sent directly to the MCW Graduate School from Educational Testing Services (ETS). Use institution code 1519. A score of 100 on the internet-based version is recommended for competitive consideration. Applicants who wish to request a waiver of the TOEFL must do so in writing via email to

  Academic Standards

On the basis of the factors listed below, individual programs will judge whether the applicant has the intellectual capability, professional commitment, research interest and experience to succeed in the program. Additional measures may be taken to determine if the applicant has the physical, communicative, and behavioral attributes necessary to complete the required course of study and research offered by the program.


Aptitude and Commitment to Research

As conveyed in a personal statement, 3 letters of recommendation, and in an interview (in person or by phone). A successful applicant will show strong motivation, passion for discovery and excitement for neuroscience research.


Experience in Research

A highly competitive applicant will have research experience in a lab, and neuroscience research is preferred.


Grade Point Average (GPA)

For basic science programs a GPA of 3.0 in science and math courses and a 3.0 overall or better. The Graduate School conducts an evaluation of foreign transcripts to determine equivalencies. Grade point average is based on a 4.0 point scale. A highly competitive applicant will have a GPA of 3.5 or higher and significant math, biology, chemistry or neuroscience coursework. 


Interpretation of Minimum Admission Requirements

The Admission Committees of individual programs evaluates the overall ability of their applicants based on these criteria and subsequently report their recommendations regarding the admission of specific applicant’s to the Dean of the Graduate School of Biomedical Sciences. The GPA and GRE scores of most matriculated students are significantly above the minimum requirements listed above, however applicants with lower scores may be offered admission in exceptional cases. In the special circumstance of recommending admission of an applicant who does not meet the minimum standards, the recommendation from the Admission Committee must be accompanied by a written explanation of the Admission Committee’s rationale for waiving the usual standards. The Dean will review that explanation along with the application and make the final admission decision.

  Technical Standards

Technical standards, distinguished from academic standards for admission to The Medical College of Wisconsin (MCW), consist of the minimum physical, cognitive, and emotional attributes required to provide reasonable assurance that a student can complete the entire course of study and participate fully in all aspects of their training. Technical standards described here are a prerequisite for admission to, and for graduation from, the Graduate School of Biomedical Sciences.


Physical Requirements

After adequate training and experience, the candidate must be capable of performing the experimental work required by the graduate program which he/she has entered. The specific requirements will vary from program to program, and within the specific research area with a program. 



A successful candidate will have strong communication skills, both in oral and written form, and a mature ability to effectively handle critique.  


Intellectual, Conceptual, Integrative and Quantitative Abilities

A candidate must have strong critical problem-solving skills in neuroscience. These skills include strong scientific rigor, honesty, understanding of the scientific method and the ability to measure, calculate, reason, analyze and synthesize. 


Behavioral and Social Attributes

A candidate must possess the emotional health, maturity and self-discipline required for full utilization of his/her intellectual ability and for successful participation in and completion of the Neuroscience Doctoral Program.



MCW may require that an applicant undergo a skills evaluation. MCW will endeavor to select and administer evaluations that accurately reflect the applicant's or student's aptitude or achievement level rather than the applicant's or student's disability.


Review of completed applications from degree seeking candidates is conducted by a faculty committee within the academic program. The outcome of the committee’s review is forwarded to the Dean as a recommendation for admission or rejection.


Offers of Admission

All offers of admission are made officially and solely by the Dean of the Graduate School of Biomedical Sciences on behalf of MCW and the academic program.  


International Applicants

International applicants must provide official documents of college/or university transcripts/mark sheets in the language of instruction and in English. If academic records do not include proof of degree award, official documents of the diplomas/certificate must also be provided. Official documents bear the original seals, stamps or signatures of the institution officials. All academic credentials are reviewed and verified by the Graduate School.  


Minority Admissions

MCW encourages the application, admission, and retention of ethnic or racial minority applicants into its graduate programs. In order to ensure that the affirmative action goals are upheld, and that all appropriate efforts are made to promote the admission of minority applicants, these applications are reviewed by both the Program Admissions Committee and, if necessary, the Admissions and Student Welfare Committee. MCW uses the same criteria as the Association of American Medical Colleges to identify minority groups.


MCW is committed to capitalizing on technological advances that create new opportunities for participation by disabled persons and will incorporate changes into its programs where feasible. A qualified disabled applicant or student shall not on the basis of his or her disability (except those which would preclude meeting the above-listed technical standards) be excluded from participation in graduate programs of the Medical College of Wisconsin.


If you have questions regarding tuition or your account, please contact the Office of Student Accounts, at (414) 955-8172 or

PhD Students

All full-time PhD students receive a full tuition remission, health insurance and stipend.

2017-2018 Stipend: $29,136.00

The following fees are covered by the tuition remission:

  • All Students Fee (Fall & Spring Semesters Only): $40.00
  • Graduate Student Association (GSA) Fee: $35.00
  • Tuition Per Credit - Fall, Spring & Summer Semesters: $1,250.00

Masters, Certificate & Non-Degree Students

Students seeking financial aid for MPH, MS or MA degree programs, visit the Financial Aid Office website.

  • Tuition Per Credit - Fall, Spring & Summer Semesters: $1,010.00
  • Master’s in Medical Physiology Tuition: $42,000/year
  • Continuation: $225.00
  • Audit - Per Class: $100.00

Current MCW Employees

Tuition Course Approval Form - Human Resources (PDF)

Late Fees

There will be a $250 late registration fee for anyone not completing registration by the date indicated on the schedule each semester. There is also a $250 late payment fee for tuition not paid on time according to the Tuition Payments policy in the Student Handbook.

Late payment fee is in addition to any late registration fee.

Faculty and Research

Neuroscience Research at the Medical College of Wisconsin is cutting-edge and highly translational. Close connections and collaborations between basic neuroscientists and clinicians make the atmosphere at MCW dynamic, disease-relevant and a rich environment for training. Our NDP students are an integral part of our research community because they receive one-on-one training from our NIH-funded faculty. If you are a prospective student looking for innovative, world-class training in a synergistic, exciting atmosphere, we invite you to explore our faculty research areas below!

Dr. Mike Lawlor
Rewriting the Whistons' Story: A Renewed Sense of Hope READ MORE ▶

Neurodegeneration and Neurotrauma

Neurodegenerative diseases including ALS, Parkinson’s Disease and Spinal Cord Injury are studied using stem cells, animal models and human tissues.

human iPSC-derived astrocytes microscopy section of the human brain cerebral organoid neurons generated from human induced pluripotent stem cells Human iPSC-Derived Motor Neurons
Xiaowen Bai, MD, PhD
Xiaowen Bai, MD, PhD

Associate Professor, Department of Anesthesiology and Physiology
My research interests are centered on the application of stem cells on disease modeling and tissue regeneration. The current major focus of the laboratory is to use utilize gain- and loss-of-function approaches to examine the novel molecular mechanisms underlying the roles of microRNAs and mitochondrial dynamics in developmental neurotoxicity in mice, and translate the findings to humans using stem cell-derived neurons and three-dimensional mini brain.

Matthew Budde, PhD
Matthew D. Budde, PhD

Associate Professor, Department of Neurosurgery
The goal of my research is to understand nervous system injury and disease and improve its diagnosis and prognosis. My primary interest is in traumatic brain and spinal cord injury and centers around magnetic resonance imaging (MRI) as a means to noninvasively detect and visualize injury. A variety of complementary and multi-disciplinary approaches are used such as preclinical models and histology, computational simulations, and MRI software and hardware development. The ultimate goal is to improve clinical care and outcomes from neurological injury.

Bonnie N. Dittel, PhD
Bonnie N. Dittel, PhD

Senior Investigator, Blood Research Institute, Blood Center of Wisconsin
The goal of my research program is to investigate the cellular and molecular mechanisms involved in the regulation of the autoimmune immune response. Broadly, we are studying how the immune system regulates inflammation associated with autoimmunity in the central nervous system (CNS) (brain and spinal cord). We are also investigating novel immune-mediated mechanisms of neurodegeneration.

Allison D. Ebert, PhD
Allison D. Ebert, PhD

Assistant Professor, Department of Cell Biology, Neurobiology and Anatomy
My lab focuses on using induced pluripotent stem cells (iPSCs) derived from patient tissue to understand the mechanisms contributing to neuronal loss. We have a particular interest in studying the neurodegenerative diseases spinal muscular atrophy, amyotrophic lateral sclerosis, and Parkinson’s disease. We use the human stem cell model system to understand the molecular basis for the disease progression and test the therapeutic benefit of small molecule treatment and gene therapy/manipulation. Finally, we validate our findings in rodent models of disease and test the therapeutic benefit of stem cell transplantation.

Balaraman Kalyanaraman, PhD
Raman Kalyanaraman, PhD

Professor and Chair, Department of Biophysics
Harry R. & Angeline E. Quadracci Professor in Parkinson’s Research
Founder, Free Radical Research Center

My research interests are centered on the application of EPR in free radical biology and understanding the role of oxidants and antioxidants in signal transduction and apoptosis.

Antje Kroner-Milsch, MD, PhD
Antje Kroner-Milsch, MD, PhD

Assistant Professor Neurosurgery (Division of Research)
Secondary appointment Microbiology and Immunology VAMC

My research explores the role of inflammation after spinal cord injury. We focus on the role macrophages play in inflammation.​ Our research aims to modify the inflammatory response in order to achieve improved outcome after spinal cord injury.

Frank A. Pintar, PhD
Frank A. Pintar, PhD

Professor, Department of Neurosurgery
Adjunct Professor, Biomedical Engineering at Marquette University
Director, VA Neuroscience Labs

Dr. Pintar's research projects are centered around the basic biomechanics of neurotrauma. Current work emphasizes the physics of concussion in military environments, and the mechanisms of spinal injury in domestic and military transportation systems. Laboratory models include experimental tissue testing and computational analysis.

Matt Scaglione, PhD
Matt Scaglione, PhD

Assistant Professor, Department of Biochemistry
The Scaglione lab investigates protein aggregation in neurodegenerative diseases. The lab has two main areas of focus, the first focuses on developing strategies to enhance neuroprotective protein quality control pathways. The second focuses on understanding how the social amoeba, Dictyostelium discoideum, naturally resists protein aggregation. Our ultimate goal is to develop therapeutics that can be used to treat neurodegenerative diseases.

Neuroimaging, tissue to brain

State-of-the-art brain imaging is used to study language, vision, hearing, learning and memory, and brain diseases including cancer.

Confocal AOSLO of Gound Squirrel Photoreceptors Combination of individual segmentation to generate radiomic profile map Cone inner segment mosaic in subject with ACHM Optos image from a subject with ocular albinism

Jeffrey R. Binder, MD
Jeffrey R. Binder, MD

Vice Chair for Research
Professor, Department of Neurology
Professor, Department of Cellular Biology, Neurobiology and Anatomy
Professor, Department of Biophysics
Director, Stroke and Neurobehavior Programs, Department of Neurology
Director, Language Imaging Laboratory, Department of Neurology
Interim Director, Functional Imaging Research Center

Our main focus is on using functional MRI to study the neurophysiological correlates of normal language processes. Though our interests range widely, the chief focus is on left temporal lobe systems associated with perceptual processes and memory stores underlying language behavior, particularly single word recognition. A second major focus of our laboratory is on development and testing of methods for presurgical functional localization of language and episodic memory systems. Our aim is to use the basic knowledge gained from fMRI studies of normal language processing to predict and prevent neuropsychological deficits in patients who must undergo surgery in sensitive brain areas.

Joseph Carroll
Joseph Carroll, PhD

Richard O. Schultz, MD / Ruth Works Professor in Ophthalmology
Professor, Department of Ophthalmology, Biophysics, and Cell Biology, Neurobiology, and Anatomy
Director, Advanced Ocular Imaging Program

My primary focus is on applying high-resolution imaging tools (adaptive optics and optical coherence tomography) to the study of the human retina.

Ted DeYoe, PhD
Edgar A. DeYoe, PhD

Professor, Department of Radiology
Professor, Department of Cell Biology, Neurobiology & Anatomy
Associate Professor, Department of Biophysics
Graduate Faculty, Programs in Cell & Developmental Biology, Neuroscience and Biophysics

In our research we seek to understand the fundamental brain processes that are responsible for our ability to see. This includes our capacities for seeing color, for accurately estimating movement, and for seeing the world as three-dimensional. It also includes more complex functions such as our ability to recognize faces.

James S. Hyde, PhD
James S. Hyde, PhD

The James S. Hyde Professor of Biophysics, Emeritus
Founder, National Biomedical EPR Center

My personal interest in EPR is to contribute to the development of EPR instrumentation and extend the ways in which existing EPR instrumentation can be used for new categories of biomedical problems.

Peter LaViolette, PhD, MS
Peter LaViolette, PhD, MS

Assistant Professor, Departments of Radiology and Biophysics
The overall goal of my lab’s research is creating and validating imaging techniques that improve both patient outcome and treatment efficacy. I primarily work with neurological diseases such as brain cancer and epilepsy in a translational setting, collaborating with clinicians for the betterment of patient care. Our brain cancer research has been focused on characterizing brain tumor cellularity and vascularity through imaging methods such as diffusion and perfusion MRI.

Shi-Jiang Li, PhD
Shi-Jiang Li, PhD

Professor, Department of Biophysics
Professor, Department of Psychiatry and Behavioral Medicine

Dr. Li's research focuses on improving and developing advanced MRI acquisition and data analysis techniques to measure brain functional and structural network organizations.

Kathleen M. Schmainda, PhD
Kathleen M. Schmainda, PhD

Robert C. Olson, MD, Professor in Radiology
Vice-chair, Imaging Research Division of Imaging Sciences

A primary focus of our laboratory is the development of MRI methods to assess brain tumor angiogenesis and invasion. The methods developed in our laboratory can provide information about the normal brain and the vascular structure of tumors, the amount of blood volume in the brain and tumor (cerebral blood volume [CBV]), and the perfusion or delivery of blood to the tumor (cerebral blood flow [CB]).

Cellular and Synaptic Communication

Neuronal Communication at the cellular level is studied using cutting-edge genetic and electrophysiological tools in order to dissect mechanisms of development and disease in the visual system, learning and memory, and addiction.

Zebrafish animal model Fluorescent reporter expression in the retina Neural progenitor development in the retina Fluorescent red nuclei and green structural proteins in the retina

Joseph Besharse, PhD
Joseph C. Besharse, PhD

Marjorie & Joseph Heil Professor in Ophthalmology
Director of Research, MCW Eye Institute

We are studying the cellular and molecular basis of circadian rhythmicity in the retina and other peripheral oscillators such as liver. The current major focus of the laboratory is to use mouse genetics to study the role of the central "clockwork" genes, Period and Clock in the retina and the role of the clock regulated genes Nocturnin and Usp2 downstream of the circadian clock.

Nashaat Gerges, PhD
Nashaat Gerges, PhD

Associate Professor, Department of Cell Biology, Neurobiology and Anatomy
The research in the laboratory is focused on elucidating the molecular and cellular mechanism of synaptic plasticity, and the associated changes in the disease states.

Iris S. Kassem, MD, PhD
Iris S. Kassem, MD, PhD

Assistant Professor of Ophthalmology
Pediatric Ophthalmology Service

By combining science with medicine, I hope to bring more knowledge to the field and discover innovative therapies to improve children’s eye care and the quality of life for our patients.

Sang Hyeong Lee, PhD
Sang Hyeong Lee, PhD

Associate Professor, Department of Pharmacology and Toxicology
The long-term goal of my research program is to understand molecular and cellular mechanisms of synaptic plasticity. Unraveling the cell biological mechanisms of how neurons control precisely the targeting, clustering, and removal of synaptic proteins into and from synapses, and how synaptic activity modulates these processes, is a fundamental goal in my lab.

Brian Link, PhD
Brian Link, PhD

Professor, Department of Cell Biology, Neurobiology and Anatomy
The overarching research goal of the Link lab is to study the cellular basis of signaling and the role in development and relationships to disease processes. We primarily use zebrafish for our studies, combining imaging based technologies and genome editing. As part of this research we have developed tools to monitor and manipulate basic cellular processes such as endocytosis, vesicle trafficking and nuclear dynamics.

Daniel M. Lipinski, MSc, DPhil
Daniel M. Lipinski, MSc, DPhil

Assistant Professor of Ophthalmology & Visual Sciences
Assistant Professor of Cell Biology, Neurobiology and Anatomy

The overarching theme of my research program is the development of broadly applicable gene-based therapeutics to preserve vision in neurodegenerative and vascular diseases affecting the retina.


Jonathan S. Marchant, MA, PhD
Jonathan S. Marchant, MA, PhD

Marcus Professor & Chair
Department of Cell Biology, Neurobiology & Anatomy (CBNA)

Dr. Marchant received training in leading laboratories that provided him with biochemical, molecular and imaging skill sets to probe the functional architecture of cell signaling events. His interests diversified to utilize these skill sets to advance a variety of projects in cellular and regenerative biology.

Qing-Song Liu, PhD
Qing-Song Liu, PhD

Associate Professor, Department of Pharmacology and Toxicology
Research in my laboratory focuses on understanding the mechanism of cocaine addiction.

Function of Neural Systems in normal and disease states

Neural circuits and pathways that underlie essential physiological functions and their misregulation in disease are studied. These include sleep disruption, breathing, chronic stress, reward systems, hearing, touch sensation and chronic pain.

Microscopy Section of the Human Brain Archeorhodopsin Trigeminal Sensory Neurons - Zebrafish Human Visual System

Carol A. Everson PhD
Carol Everson, PhD

Professor, Department of Medicine
Our mission is to determine how the restriction of sleep, a basic biological requirement, affects the functions of cells, processes and systems in ways that increase the risk of disease, such as cardiovascular disease, osteoporosis, and cancer. Areas of study include mediation of cell injury and inflammatory processes by sleep, abnormalities in blood cell production and bone metabolism resulting from sleep restriction, and changes to brain injury status by sleep augmentation and sleep restriction.

 Hubert V. Forster, PhD
Hubert V. Forster, Ph.D

Professor, Department of Physiology
Research in my laboratory is primarily concerned with mechanisms regulating breathing. Our studies relate to disease conditions of central and obstructive sleep apnea, Sudden Infant Death syndrome, congenital central alveolar hypoventilation, and traumatic brainstem injury.

David R. Harder PhD
David R. Harder, PhD

Kohler Co. Professor in Cardiovascular Research and Associate Dean for Research
Professor of Physiology, Medicine and Pediatrics
Director for Mentoring – Clinical and Translational Science Institute

The major focus of our laboratory is to understand mechanisms regulating the cerebral circulation under physiologic and pathophysiologic conditions. Studies in the laboratory are focused around the astrocyte as both a homeostatic sensor and an active regulator that can affect and protect both the cerebral vasculature and neurons.

Cecilia J. Hillard, PhD
Cecilia J. Hillard, PhD

Professor, Department of Pharmacology and Toxicology
Director of the Neuroscience Research Center

A long-standing interest of our laboratory is the study of the mechanisms by which the cannabinoids affect the function of the brain.

Matthew R. Hodges , PhD
Matthew R. Hodges , PhD

Assistant Professor, Department of Physiology
Our research is focused on understanding the roles of serotonin-producing neurons in controlling breathing and body temperature in health and disease. We use molecular genetics and cellular electrophysiologic techniques to identify genes and/or proteins that underlie pH sensing in the brain. We also study transgenic, mutant, and knockout rats as models of human diseases including SIDS, bronchopulmonary dysplasia (BPD) and epilepsy.

John R Kirby, PhD
John R. Kirby, PhD

Chair, Department of Microbiology and Immunology

Walter Schroeder Professor in Microbiology and Immunology

Dr. Kirby’s major areas of research focus on signal transduction in diverse bacteria ranging from soil dwelling spore formers (Bacillus subtilis and Myxococcus xanthus) to biofilm forming pathogens, to microbial communities in the gut. Dr. Kirby is actively investigating interactions between M. xanthus and B. subtilis as a model for predator-prey interactions in vivo, primarily to assess the role of production of specialized metabolites, similar to antibiotics, on both sides of the predator-prey equation. Additionally, he has been examining the role of xenobiotics (antipsychotics, antihypertensives and antibiotics) for their capacity to disrupt the gut microbiota with deleterious consequences on metabolism.

Michael W. Lawlor, MD, PhD
Michael W. Lawlor, MD, PhD

Assistant Professor of Pathology
Director, Pediatric Pathology Neuromuscular Laboratory

My work focuses extensively on testing treatments for XLMTM in animal models, and we now are in the process of translating this knowledge for the planning of human clinical trials. We continue to participate in a number of projects including:

  • Pediatric muscle disease (X-linked myotubular myopathy)
  • Preclinical trials of novel therapeutic agents
  • Skeletal muscle pathology
  • Skeletal muscle physiology
  • In vitro models of skeletal muscle function and disease


Christopher M. Olsen, PhD
Christopher M. Olsen, PhD

Assistant Professor, Department of Pharmacology and Toxicology and Neuroscience Research Center
The Olsen lab studies the neural underpinnings of behavior motivated by natural rewards and drugs of abuse. We use transgenic approaches to identify and manipulate neurons that are strongly engaged during specific behaviors such as cocaine seeking. In collaboration with faculty in the Department of Neurosurgery, we also study how mild brain injury can impact drug seeking and emotional behavior.

Danny A. Riley, PhD
Danny A. Riley, PhD

Professor, Department of Cell Biology, Neurobiology and Anatomy
My research focuses on understanding vibration injury of neural and vascular tissues from hand-held powered tools and the effects of actual and simulated spaceflight unloading on neuromuscular structure and function.

Cheryl Stucky, PhD
Cheryl Stucky, PhD

Professor, Department of Cell Biology, Neurobiology and Anatomy
Director, Neuroscience Doctoral Program

The Stucky Lab has made key contributions to the pain field’s understanding of how ion channels on pain-sensing neurons contribute to pain and touch sensation. An exciting current direction is identifying mechanisms underlying pain in damaged skin, by examining bidirectional signaling between keratinocytes and sensory neurons. Another major focus is translational models of chronic pain including inflammation, nerve injury, sickle cell disease and Fabry Disease that are associated with devastating pain.

Margaret Wong-Riley, PhD
Margaret Wong-Riley, PhD

Professor, Department of Cell Biology, Neurobiology and Anatomy
Two areas of research: (1) We are probing for the molecular mechanisms underlying a tight coupling between neuronal activity and energy metabolism that are the basis for health and neurodegenerative diseases; and (2) we are uncovering the metabolic, neurochemical, ventilatory and electrophysiological bases of a critical period of postnatal respiratory development that may have significant implications for Sudden Infant Death Syndrome (SIDS).

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Neuroscience Doctoral Program Alumni

Sheila Baker, PhD
Assistant Professor of Biochemistry
Assistant Professor of Ophthalmology and Visual Sciences
University of Iowa
PhD Mentor: Joseph Besharse, PhD

Alicia Barr, PhD
Postdoctoral Research in Lisdahl Lab, Psychology
University of Wisconsin Madison
PhD Mentor: Alan Bloom, PhD

Diane Bishop (Tait), PhD
Spark Therapeutics
PhD Mentor: Jay Neitz, PhD

Julie Brefczynski-Lewis, PhD
Research Assistant Professor, Radiology
West Virginia University
PhD Mentor: Edgar DeYoe, PhD

Brian Clark, PhD
Postdoctoral Research Fellow
The Solomon H. Snyder Department of Neuroscience
Johns Hopkins University
PhD Mentor: Brian Link, PhD

Brian Curry, PhD, MBA
President & Chief Executive Officer at Simply 1Life Inc
Adjunct Faculty
Carroll University, Concordia University
PhD Mentor: Danny Riley, PhD

Joseph Fogerty, PhD
Postdoctoral Fellow, Department of Ophthalmology
Cleveland Clinic
PhD Mentor: Joseph Besharse, PhD

Wendy Huddleston, PT, PhD
Associate Professor
University of Wisconsin Milwaukee
PhD Mentor: Edgar DeYoe, PhD

Christine (Insinna) Kettenhofen, PhD
Postdoctoral Fellow (CRTA)
Center for Cancer Research, National Cancer Institute
PhD Mentor: Joseph Besharse, PhD

Sivesh Pillay, PhD
Postdoctoral Fellow, Anesthesiology
Medical College of Wisconsin
PhD Mentor: Tony Hudetz, PhD

Michael A. Pizzi, DO, PhD
Medical Director of the Neuroscience Intensive Care Unit
Assistant Professor of Neurology
University of Florida - Jacksonville
PhD Mentor: Maria Crowe, PhD

Alexander Puckett, PhD
Postdoctoral Researcher
The University of Queensland, Australia
PhD Mentor: Edgar DeYoe, PhD

Daniel Roberson, PhD
Watertown Regional Medical Center
PhD Mentor: Jay Neitz, PhD

Andrew Salzwedel, PhD
Project Scientist
PhD Mentor: Edgar DeYoe, PhD

Leah Shriver, PhD
Assistant Professor, Departments of Chemistry and Biology
University of Akron
Co-Founder, Therakinetix
PhD Mentor: Bonnie Dittel, PhD

Benjamin Stengel, PhD
Project Manager
Epic Software
PhD Mentor: Jeffrey Binder, MD

Daniel Vilceanu, MD, PhD
Assistant Professor of Clinical Anesthesiology
School of Medicine, University of Missouri Health
PhD Mentor: Cheryl Stucky, PhD

Melissa Wilk, PhD
Postdoctoral Researcher
HudsonAlpha Institute for Biotechnology
PhD Mentor: Joseph Carroll, PhD

Graduate School of Biomedical Sciences General CAMPUS CONTACT INFORMATION

Mailing Address:
MCW Graduate School
8701 Watertown Plank Road
Milwaukee, WI 53226

(414) 955-8218
(414) 955-6555 (fax)

Neuroscience Doctoral Program

(414) 955-4402
(414) 955-6555 (fax)
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