Faculty
BIOCHEMISTRY
Professor and Chairman
John A. Corbett, PhD
PhD – Utah State, 1990
Professor
Nancy M. Dahms, PhD
PhD – Johns Hopkins University, 1985
Albert W. Girotti, PhD
PhD – University of Massachusetts, 1965
Owen W. Griffith , PhD
PhD – The Rockefeller University, 1974
Vaughn E. Jackson, PhD
PhD – University of Iowa, 1972
Jung-Ja P. Kim, PhD
PhD – Cornell University, 1969
Ravi P. Misra, PhD
Dean of the Graduate School of Biomedical Sciences
PhD – New York University-School of Medicine
Sackler Institute, 1988
Sally S. Twining, PhD
PhD – Ohio State University, 1976
Brian F. Volkman, PhD
PhD – University of California-Berkley, 1994
Associate Professor
Bassam T. Wakim, PhD
PhD – University of Freiburg, Germany, 1980
Assistant Professor
Jianhua Fu, PhD
PhD – University of Pittsburgh, 1994
Witold Korytowski, PhD
PhD – Jagiellonian University, Poland, 1983
Evgenii Kovrigin, PhD
PhD – Institute of Protein Chemistry, Russia, 1999
Jong-In Park, PhD
PhD – University of New South Wales, Australia, 2000
Francis C. Peterson, PhD
PhD – Ohio State University-Columbus 1997
Shama Mirza, PhD
Indian Institute of Chemical Technology, India, 2004
Secondary Appointments
Executive Vice President for Research/Associate Dean of Research
Gilbert C. White II, MD
MD – University of North Carolina at Chapel Hill
School of Medicine, 1971
Professor
John E. Baker, PhD
PhD – St. Thomas Medical School, England, 1984
Andrew Chan, PhD
PhD – University of London (Institute of Cancer Research), 1988
Assistant Professor
Victor Drover, PhD
PhD – University of Alberta, 2002
BIOPHYSICS
Professor and Chairman
Balaraman Kalyanaraman, PhD
PhD – University of Alabama, 1978
Professor
Jimmy B. Feix, PhD
PhD – University of Kentucky, 1981
Neil Hogg, PhD
Assistant Dean of Students for the Graduate School of Biomedical Sciences
PhD – University of Essex, 1992
James S. Hyde, PhD
Director of the National Biomedical EPR Center
PhD – Massachusetts Institute of Technology, 1959
Shi-Jiang Li, PhD
Director of the Center for Imaging Research and Animal Imaging Program
PhD – Ohio State University, 1985
Witold Subczynski, PhD
PhD – Lomonosov Moscow State University, 1976
Associate Professor
William E. Antholine, PhD
PhD – Iowa State University, 1971
Brian Bennett, PhD
PhD – University of Sussex, 1994
Andrzej Jesmanowicz, PhD
PhD – Copernicus University, 1975
Candice Klug, PhD
PhD – Medical College of Wisconsin, 1999
Jeannette M. Vasquez-Vivar, PhD
PhD – Instituto de Quimica Universidade de Sao Paul, 1992
Ming Zhao, PhD
PhD – University of Cambridge, 2000
Secondary Appointments
Professor
Jeffery R. Binder, MD
MD – University of Nebraska College of Medicine, 1986
Edgar A. DeYoe, PhD
PhD – University of Rochester, 1983
Andrew S. Greene, PhD
Director of Biotechnology and Bioengineering Center
PhD – Johns Hopkins University, 1985
Anthony G. Hudetz, PhD
PhD – Semmelweis Orvostudomanyi Egyetem Budapest, 1985
Kathleen M. Schmainda, PhD
PhD – Harvard University, 1993
Reza Shaker, MD
Joseph E. Geenen Professor and Chief
Senior Associate Dean for Clinical and Translational Research
MD – University of Tehran, Pahlavi Medical School, 1975
Associate Professor
Robert W. Prost, PhD
PhD – Medical College of Wisconsin, 1999
CELL BIOLOGY, NEUROBIOLOGY & ANATOMY
Professor and Chairman
Joseph C. Besharse, PhD
Marvin Wagner Professor in Cell Biology, Neurobiology and Anatomy
PhD – Southern Illinois University at Carbondale, 1973
Professor
Stephen A. Duncan, D.Phil.
Marcus Professor in Human and Molecular Genetics
D.Phil. – Oxford University, 1992
Gary L. Kolesari, MD, PhD
MD – Medical College of Wisconsin, 1977
PhD – Medical College of Wisconsin, 1976
John W. Lough, PhD
PhD – Washington University, 1975
Danny A. Riley, PhD
PhD – University of Wisconsin, 1972
Cheryl L. Stucky, PhD
PhD – University of Minnesota, 1995
Margaret T. Wong-Riley, PhD
PhD – Stanford University, 1970
Associate Professor
Lisa A. Cirillo, PhD
PhD – University of Illinois-Urbana-Champaign, 1996
Brian A. Link, PhD
PhD – Oregon Health Sciences University, 1997
Duska J. Sidjanin, PhD
PhD – Northern Illinois University, 1995
Assistant Professor
Michele A. Battle, PhD
PhD – Michigan State University, 2002
Nashaat Z. Gerges, PhD
PhD – University of Houston, 2002
Secondary Appointments
Professor
Janice M. Burke, PhD
Marjorie & Joseph Heil Professor in Ophthalmology
PhD – University of Massachusetts, 1972
Carol A. Everson, PhD
PhD – University of Chicago, 1987
Peter J. Newman, PhD
PhD – St. Louis University, 1983
Elena Semina, PhD
PhD – National Center of Medical Genetics
Russian Academy, 1993
Associate Professor
Ramani Ramchandran, PhD
PhD – Medical College of Georgia School of Medicine, 1997
Assistant Professor
Joseph J. Carroll, PhD
PhD – Medical College of Wisconsin, 2002
Maria J. Crowe, PhD
PhD – University of Cincinnati, 1992
Vivian Lee, PhD
PhD – Oregon Health Sciences University, 2001
Alan N. Mayer, MD, PhD
MD – Cornell University Medical School, 1995
PhD – Cornell University Medical School, 1994
Katherine Shim, PhD
PhD – Stanford University, 2001
George Wilkinson, PhD
PhD – University of California-San Francisco, 1997
INSTITUTE FOR HEALTH AND SOCIETY
Professor and Director
Cheryl A. Maurana, PhD
Senior Associate Dean for Public and Community Health
PhD – Purdue University, 1976
Professor and Director – Center for Bioethics and Medical Humanities
Arthur R. Derse, MD, JD
Julia and David Uihlein Professor in Medical Humanities
MD – University of Wisconsin Medical School, 1980
JD – University of Wisconsin Law School, 1987
Professor and Director – Biostatistics
John P. Klein, PhD
PhD – University of Missouri-Columbia, 1980
Professor
Jane Morley Kotchen, MD, MPH
MD – Case Western Reserve University, 1964
MPH – University of North Carolina-Chapel Hill, 1966
Purushottam W. Laud, PhD
PhD – University of Missouri-Columbia, 1977
C. Oren Renick, JD, MPH
JD – Mississippi College, 1983
MPH – Tulane University, 1974
David Wyatt Seal, PhD
PhD – University of New Mexico, 1994
Mei-Jie Zhang, PhD
PhD – Florida State University, 1991
Associate Professor
Brent R. Logan, PhD
PhD – Northwestern University, 2001
Thomas May, PhD
PhD – Bowling Green State University, 1994
Ryan B. Spellecy, PhD
PhD – University of Utah, 2002
Aniko Szabo, PhD
PhD – University of Tennessee, Memphis, College of Medicine, 1998
Tao Wang, PhD
PhD – North Carolina State University, Raleigh, 2001
Assistant Professor
Kwang Woo Ahn, PhD
PhD – University of Iowa, 2008
Ruta Brazauskas, PhD
PhD – Medical College of Wisconsin, 2003
Jennifer Le-Rademacher, PhD
PhD – University of Georgia, 2008
Jessica Pruszynski, PhD
PhD – Baylor University, 2010
Sergey S. Tarima, PhD
PhD – University of Kentucky, 2005
Associate Clinical Professor
William W. Greaves, MD, MSPH
MD – University of Illinois College of Medicine, 1975
MSPH – University of Utah, 1980
Cynthiane J. Morgenweck, MD, MA
MD – Medical College of Wisconsin, 1977
MA – Medical College of Wisconsin, 1999
Associate Adjunct Professor
Michael D. Akers, PhD
PhD – University of Mississippi, 1988
Dean B. McFarlin, PhD
PhD – State University of New York at Buffalo, 1986
Matthew Wynia, MD, MPH
MD – Oregon Health Sciences University School of Medicine, 1990
MPH – Harvard University, 1997
Assistant Clinical Professor
Ross L. Clay, MD, MPH
MD – University of Minnesota Medical School, 1983
MPH – University of Minnesota School of Public Health, 1989
Jay A. Gold, MD, JD, MPH
MD – Texas A&M University College of Medicine, 1986
JD – New York University, 1974
MPH – Harvard University, 1977
Robert L. Goldberg, MD, FACOEM
MD – Jefferson Medical College, 1976
Leslie F. Martin, MD, MPH
MD – University of Lille Medical School, 1985
MPH – Columbia University School of Public Health, 1991
Assistant Adjunct Professor
J. Todd Ferguson, PhD, MA
PhD – Purdue University, 2004
MA – Purdue University, 2001
Nancy J. Kreuser, RN, PhD
PhD – University of Wisconsin-Milwaukee, 1998
Kristin E. Schleiter, JD
JD – Loyola University Chicago School of Law, 2007
Julia A. Uihlein, MA
MA – Medical College of Wisconsin, 1998
Julie A. Willems Van Dijk, PhD, MSN
PhD – University of Wisconsin-Milwaukee, 2008
MSN – University of Wisconsin-Oshkosh, 1991
Secondary Appointments
Professor
Syed M. Ahmed, MD, MPH, DrPH, FAAFP
Associate Dean for Public and Community Health
MD – Sir Salimullah Medical College, Dhaka University, 1979
MPH – University of Texas-Houston, 1984
DrPH – University of Texas-Houston, 1990
Bruce M. Camitta, MD
MD – Johns Hopkins University School of Medicine, 1966
David C. Clark, PhD
PhD – University of Chicago, 1981
Marc H. Gorelick, MD, MSCE
MD – Duke University School of Medicine, 1987
MSCE – University of Pennsylvania, 1996
David D. Gutterman, MD
Northwestern Mutual Professor in Cardiology
MD – University of North Carolina at Chapel Hill School of Medicine, 1980
Stephen H. Hargarten, MD, MPH
Associate Dean for Global Health
MD – Medical College of Wisconsin, 1975
MPH – Johns Hopkins School of Public Health, 1984
Peter L. Havens, MD
MD – University of Missouri Kansas City
School of Medicine, 1980
MS – Harvard School of Public Health, 1987
Raymond G. Hoffmann, PhD
PhD – Johns Hopkins University, 1978
Richard L. Holloway, PhD
PhD – Syracuse University, 1976
Peter M. Layde, MD, MSc
MD – University of Wisconsin Medical School, 1976
MSc – London School of Hygiene and Tropical Medicine, 1980
Timothy L. McAuliffe, PhD
PhD – University of California, 1981
John R. Meurer, MD, MBA
MD – University of Wisconsin Medical School, 1986
MBA – Northwestern University, 1991
Ann B. Nattinger, MD, MPH
Lady Riders Breast Cancer Research Professor
MD – University of Illinois College of Medicine, 1983
MPH – University of Rochester School of Medicine & Dentistry, 1988
Steven D. Pinkerton, PhD
PhD – University of California-Los Angeles, 1995
Marilyn M. Schapira, MD, MPH
MD – University of Michigan Medical School, 1986
MPH – University of Michigan School of Public Health, 1986
Earnestine Willis, MD, MPH
MD – Harvard Medical School, 1976
MPH – Harvard School of Public Health, 1977
Associate Professor
David C. Brousseau, MD, MS
MD – University of Virginia School of Medicine, 1993
MS – Brown University, 2000
Laura Cassidy, PhD, MS
PhD – University of Pittsburgh, 2000
MS – University of Pittsburgh, 1993
Christine E. Cronk, ScD
ScD – Harvard School of Public Health, 1980
MS – Harvard School of Public Health, 1977
Mary Eapen, MBBS, MS
MBBS – Ahmadu Bello University, 1987
MS – University of Minnesota, 2000
E. Brooke Lerner, PhD
PhD – State University of New York-Buffalo, 2001
Linda N. Meurer, MD, MPH
MD – Wayne State University School of Medicine, 1989
MPH – University of Minnesota School of Public Health, 1994
Jeffrey A. Morzinski, PhD
PhD – University of Wisconsin-Milwaukee, 1999
Kenneth G. Schellhase, MD, MPH
MD – University of Pennsylvania School of Medicine, 1994
MPH – University of Washington School of Public Health, 1999
Mary Lou Taylor, PhD
PhD – Kent State University, 1987
Grzegorz W. Telega, MD
MD – N Copernicus Academy of Medicine, 1989
Jerome V. Van Ruiswyk, MD
MD – University of Wisconsin Medical School, 1985
Assistant Professor
Suzanne N. Brixey, MD
MD – University of Virginia School of Medicine, 1998
Elaine M. Drew, PhD
PhD – University of Kentucky, 2004
Michael H. Farrell, MD
MD – St. Louis University, School of Medicine, 1993
Sheri Johnson, PhD
PhD – Boston University, 1993
David Nelson, PhD, MA
PhD – University of Tennessee, Knoxville, 2000
MA – University of Northern Colorado, 1995
Joan M. Neuner, MD, MPH
MD – University of Wisconsin Medical School, 1994
MPH – Harvard School of Public Health, 2000
Emmanuel M. Ngui, DrPH, MSc
DrPH – University of North Carolina at Chapel Hill School of Public Health, 2003
MSc – Eastern University, 1995
Andrew E. Petroll, MD, MS
MD – University of Wisconsin, Madison, 2001
MS – Medical College of Wisconsin, 2008
Alan Wells, PhD, MPH
PhD – University of Texas School of Public Health, 2003
MPH – University of Texas School of Public Health, 1997
Associate Clinical Professor
N. Timothy Lynch, PhD
PhD – University of Wisconsin Medical School, 1973
MICROBIOLOGY & MOLECULAR GENETICS
Professor and Chairman
Paula Traktman, PhD
Walter Schroeder Chair in Microbiology
PhD – Massachusetts Institute of Technology, 1981
Professor
Joseph T. Barbieri, PhD
PhD – University of Massachusetts, 1980
Dara W. Frank, PhD
Director of the Center for Infectious Disease Research
PhD – University of Texas-Austin, 1984
Associate Professor
Michael B. Dwinell, PhD
PhD – University of Wisconsin, 1996
Amy W. Hudson, PhD
PhD – Harvard University, 1995
Roy M. Long, PhD
Assistant Dean of Recruitment for the Graduate School of Biomedical Sciences
PhD – Pennsylvania State University College of Medicine, 1994
Mark T. McNally, PhD
PhD – State University of New York at Buffalo, 1987
Thomas C. Zahrt, PhD
PhD – University of Illinois-Urbana-Champaign, 1997
Assistant Professor
William T. Jackson, PhD
PhD – University of California-Berkeley, 1999
Christopher J. Kristich, PhD
PhD – University of Illinois –Urbana-Champaign, 2002
Vera Tarakanova, PhD
PhD – St. Louis University, 2004
Scott S. Terhune, PhD
PhD – Northwestern University Medical School, 2000
Secondary Appointments
Professor
Jenifer Coburn, PhD
PhD – Tufts University Sackler School of Graduate Biomedical Sciences, 1991
William R. Drobyski, MD
MD – University of Rochester School of Medicine and Dentistry, 1983
Bryon D. Johnson, PhD
PhD – University of Health Sciences/The Chicago Medical School, 1989
John M. Routes, MD
MD – Indiana University School of Medicine, 1981
Andrey B. Sorokin, PhD
PhD – Institute of Cytology Academy of Sciences, 1980
Associate Professor
Soumitra Ghosh, PhD
PhD – Oxford University, 1992
Jack Gorski, PhD
PhD – University of Cincinnati, 1975
Mitchell H. Grayson, MD
MD – University of Chicago/Pritzker School of Medicine, 1993
Martin J. Hessner, PhD
PhD – University of Wisconsin-Milwaukee, 1990
Subramaniam Malarkannan, PhD
PhD – Madurai Kamaraj University, 1992
Nita H. Salzman, MD, PhD
MD – New York University School of Medicine, 1990
PhD – New York University School of Medicine, 1989
Calvin B. Williams, MD, PhD
MD – University of California Irvine School of Medicine, 1991
PhD – University of California Irvine School of Medicine, 1991
Assistant Professor
Stephen B. Gauld, PhD
PhD – University of Glasgow, 2000
James Verbsky, MD, PhD
MD – Washington University School of Medicine, 2000
PhD – Washington University School of Medicine, 2000
Assistant Adjunct Professor
Bonnie Dittel, PhD
PhD – University of Minnesota, 1994
Laurent Malherbe, PhD
PhD – University of Nice, 2001
Debra Newman, PhD
PhD – Marquette University, 1989
Demin Wang, PhD
PhD – University of Tennessee, Memphis College of Medicine, 1995
PHARMACOLOGY & TOXICOLOGY
Professor and Chairman
William B. Campbell, PhD
PhD – University of Texas Health Science Center, 1974
Professor
John A. Auchampach, PhD
PhD – Medical College of Wisconsin, 1994
Alan S. Bloom, PhD
PhD – University of Louisville, 1973
Guan Chen, MD, PhD
MD – Bengbu Medical College, 1982
PhD – University of Heidelberg, Germany, 1990
Garrett J. Gross, PhD
PhD – University of Utah, 1971
Cecilia J. Hillard, PhD
Director of Neuroscience Research Center
PhD – Medical College of Wisconsin, 1983
John D. Imig, PhD
PhD – University of Louisville, 1990
Charles R. Myers, PhD
PhD – University of Wisconsin-Milwaukee, 1987
Peter J. Newman, PhD
PhD – St. Louis University, 1983
Kasem Nithipatikom, PhD
PhD – Marquette University, 1985
Carol L. Williams, PhD
PhD – University of Minnesota, 1983
Associate Professor
Kathryn M. Gauthier, PhD
PhD – Medical College of Wisconsin, 1998
Sandra L. Pfister, PhD
PhD – University of Texas Health Science Center, 1984
Assistant Professor
Sang Hyeong Lee, PhD
PhD – Gyeongsang National University, 1998
Qing-song Liu, PhD
PhD – Academy of Military Medical Sciences, 1994
Xiao-Mei Qi, MD
MD – Bengbu Medical School, 1982
Secondary Appointments
Professor
John E. Baker, PhD
PhD – St. Thomas Medical School, England, 1984
Janice M. Burke, PhD
Marjorie & Joseph Heil Professor in Ophthalmology
PhD – University of Massachusetts, 1972
David D. Gutterman, MD
Northwestern Mutual Professor in Cardiology
MD – University of North Carolina at Chapel Hill
School of Medicine, 1980
Ronald N. Hines, PhD
PhD – University of Texas Southwestern
Medical Center at Dallas, 1976
Judy R. Kersten, MD
MD – University of Wisconsin Medical School, 1988
Wai Meng Kwok, PhD
PhD – University of Illinois-Urbana-Champaign, 1990
D. Gail McCarver, MD
MD – University of Tennessee –
Memphis College of Medicine, 1978
Marilyn P. Merker, PhD
PhD – Yale University, 1983
Kirkwood A. Pritchard, PhD
PhD – Ohio State University, 1983
David L. Roerig, PhD
PhD – Kansas State University, 1970
David C. Warltier, MD, PhD
The John P. Kampine Professor in Anesthesiology
MD – Medical College of Wisconsin, 1982
PhD – Medical College of Wisconsin, 1976
Gilbert C. White II, MD
MD – University of North Carolina at Chapel Hill
School of Medicine, 1971
Associate Professor
Constantine Sarantopoulos, MD, PhD
MD – University of Athens, Greece, 1985
PhD – University of Athens, Greece, 1992
Assistant Professor
Martin Bienengraeber, PhD
PhD – University of Munich, 1998
Qing Miao, PhD
PhD – Medical University of South Carolina, 2001
PhD – Dalian Institute of Chemical Physics, Chinese Academy of Sciences, 1995
Debra Newman, PhD
PhD – Marquette University, 1989
Phillip F. Pratt, PhD
PhD – Medical College of Wisconsin, 1998
Daisy Sahoo, PhD
PhD – University of Alberta, 2000
Michael E. Widlansky, MD
MD – University of Michigan, 1999
Assistant Adjunct Professor
Magdalena Wodnicka, PhD
PhD – University of North Carolina at Chapel Hill, 1997
PHYSIOLOGY
Professor and Chairman
Allen W. Cowley, PhD
James J. Smith & Catherine Welsh Smith Professor in Physiology
Harry & Gertrude Hack Term Professor in Physiology
Director of the Cardiovascular Center
PhD – Hahnemann University, 1968
Professor
Daniel A. Beard, PhD
PhD – University of Washington, 1997
Hubert V. Forster, PhD
PhD – University of Wisconsin, 1968
Andrew S. Greene, PhD
Director of Biotechnology and
Bioengineering Center
PhD – Johns Hopkins University, 1985
David R. Harder, PhD
Associate Dean for Research Mentoring
Kohler Co. Professor in Cardiovascular Research
PhD – Medical College of Wisconsin, 1976
Howard J. Jacob, PhD
Warren P. Knowles Professor in Human and Molecular Genetics
Director of the Human and Molecular Genetics Center
PhD – University of Iowa, 1989
Julian H. Lombard, PhD
PhD – Medical College of Wisconsin, 1974
David L. Mattson, PhD
PhD – Medical College of Wisconsin, 1990
Michael Olivier, PhD
PhD – Cornell University, 1997
William J. Stekiel, PhD
PhD – Johns Hopkins University, 1957
Associate Professor
Ranjan K. Dash, PhD
PhD – Indian Institute of Technology, 1998
Debebe Gebremedhin, PhD
PhD – Semmelweis Orvostudományi Egyetem, Budapest, 1988
Matthew R. Hodges, PhD
PhD – Medical College of Wisconsin, 2004
Mingyu Liang, PhD
PhD – Mayo Graduate School, 1999
Assistant Professor
Melinda R. Dwinell, PhD
PhD – University of Wisconsin, 1996
Diane H. Munzenmaier, PhD
PhD – Medical College of Wisconsin, 1995
Carol M. Quinn, MD, PhD
MD – Universidad de Murcia, 1995
PhD – Universidad de Murcia, 1999
Paul O’Connor, PhD
PhD – Monash University, 2005
Andrew K. Tryba, PhD
PhD – Case Western Reserve University, 2000
Simon N. Twigger, PhD
PhD – University of Nottingham, 1994
Tetsuro Wakatsuki, PhD
PhD – Washington University School of Medicine, 1999
Hartmut Weiler, PhD
PhD – Technische Hochschule Darmstadt, 1989
Secondary Appointments
Professor
Zeljko J. Bosnjak, PhD
PhD – Medical College of Wisconsin, 1979
Philip S. Clifford, PhD
Associate Dean for Postdoctoral Education
PhD – Medical College of Wisconsin, 1985
Michael J. Dunn, MD
Distinguished Professor and Dean Emeritus
MD – Marquette University School of Medicine, 1962
Elizabeth R. Jacobs, MD
MD – University of Kansas School of Medicine, 1977
Hershel Raff, PhD
PhD – Johns Hopkins University, 1981
Jeanne L. Seagard, PhD
PhD – Medical College of Wisconsin, 1978
David F. Stowe, MD, PhD
MD – Medical College of Wisconsin, 1983
PhD – Michigan State University, 1974
Associate Professor
Caron Dean-Bernhoft, PhD
PhD – Birmingham University, 1984
Jozef Lazar, MD, PhD
MD – Comenius University, 1980
PhD – Comenius University, 1987
Anna Stadnicka, PhD
PhD – Jagiellonian University, Krakow, Poland, 1975
Tao Wang, PhD
PhD – North Carolina State University, Raleigh, 2001
OFFICE OF TECHNOLOGY DEVELOPMENT
Vice President Technology Development
Joseph O. Hill, PhD
PhD – West Virginia University, 1977